Background: Richard Charles "Dick" Lewontin (born March 29, 1929) is an American evolutionary biologist, mathematician, geneticist, and social commentator. A leader in developing the mathematical basis of population genetics and evolutionary theory, he pioneered the application of techniques from molecular biology, such as gel electrophoresis, to questions of genetic variation and evolution. In a pair of seminal 1966 papers co-authored with J.L. Hubby in the journal Genetics, Lewontin helped set the stage for the modern field of molecular evolution. In 1979 he and Stephen Jay Gould introduced the term "spandrel" into evolutionary theory.
Context: Lewontin has worked in both theoretical and experimental population genetics. A hallmark of his work has been an interest in new technology. He was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci). In 1960 he and Ken-Ichi Kojima were the first population geneticists to give the equations for change of haplotype frequencies with interacting natural selection at two loci. This set off a wave of theoretical work on two-locus selection in the 1960s and 1970s. Their paper gave a theoretical derivation of the equilibria expected, and also investigated the dynamics of the model by computer iteration. Lewontin later introduced the D' measure of linkage disequilibrium. (He also introduced the term "linkage disequilibrium", about which many population geneticists have been unenthusiastic.)  In 1966, he and Jack Hubby published a paper that revolutionized population genetics. They used protein gel electrophoresis to survey dozens of loci in the fruit fly Drosophila pseudoobscura, and reported that a large fraction of the loci were polymorphic, and that at the average locus there was about a 15% chance that the individual was heterozygous. (Harry Harris reported similar results for humans at about the same time.) Previous work with gel electrophoresis had been reports of variation in single loci and did not give any sense of how common variation was.  Lewontin and Hubby's paper also discussed the possible explanation of the high levels of variability by either balancing selection or neutral mutation. Although they did not commit themselves to advocating neutrality, this was the first clear statement of the neutral theory for levels of variability within species. Lewontin and Hubby's paper had great impact--the discovery of high levels of molecular variability gave population geneticists ample material to work on, and gave them access to variation at single loci. The possible theoretical explanations of this rampant polymorphism became the focus of most population genetics work thereafter. Martin Kreitman was later to do a pioneering survey of population-level variability in DNA sequences while a Ph.D. student in Lewontin's lab.
Question: What else did he accomplish?
Answer: technology. He was the first person to do a computer simulation of the behavior of a single gene locus (previous simulation work having been of models with multiple loci).

Problem: Background: Khloe Alexandra Kardashian was born in Los Angeles, California on June 27, 1984, to Robert, an attorney, and Kris (nee Houghton), a homemaker. She has two older sisters Kourtney and Kim, and a younger brother Rob. Their mother is of Dutch, English, Irish and Scottish ancestry, while their father was a third-generation Armenian American. After their parents divorced in 1991, Houghton remarried the 1976 Summer Olympics decathlon winner Caitlyn Jenner (then Bruce) in 1991.
Context: In 2001, Kardashian suffered a traumatic brain injury in a car accident. She went through the windscreen and suffered a severe concussion, causing long-term memory loss.  On September 27, 2009, Kardashian married NBA basketball player Lamar Odom, who was a member of the Los Angeles Lakers at the time. The couple were married exactly one month after they met at a party for Odom's teammate Metta World Peace. Following her marriage, Kardashian removed her middle name to include her married surname, becoming Khloe Kardashian Odom. Kardashian adopted a pet boxer named Bernard "BHops" Hopkins, after the famous boxer.  On December 13, 2013, after months of speculated separation, Kardashian filed for divorce from Odom and for legal restoration of her last name. Divorce papers were signed by both parties in July 2015; however, the divorce had yet to receive final approval from a judge. In October 2015, Odom was hospitalized after being found unconscious in a Nevada brothel, and was in a coma for four days; as he lay in a hospital, Kardashian withdrew her pending divorce petition. In an interview with People Magazine, Kardashian confirmed that they had not reconciled and the divorce had been withdrawn so that she might make medical decisions on Odom's behalf. Kardashian and Odom's divorce was finalized in December 2016.  Kardashian is currently in a relationship with basketball player Tristan Thompson. The couple resides in Cleveland, Ohio. In December 2017, she announced she was expecting their first child together. In the March 2018 she revealed she will be having a girl.  Kardashian supports the recognization of the Armenian Genocide and has visited the memorial of the victims in Yerevan, Armenia. In April 2015, Kardashian accompanied her sister, Kim, and brother-in-law, Kanye West, to Jerusalem for the baptism of her niece, North, in the Armenian Apostolic Church at the Cathedral of St. James. She was named North's godmother during the ceremony.
Question: What is significant about Khloe's personal life ?
Answer:
On December 13, 2013, after months of speculated separation, Kardashian filed for divorce from Odom and for legal restoration of her last name.