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Bourke's father, Frank, was serving in the RAAF and on leave in Melbourne when he played a solitary game for Richmond in 1943. A tall (193 cm) and lean (85 kg) full forward with an excellent reputation in country football, Frank returned to the city after the war and resume his playing career with Richmond. In the opening weeks of the 1946 season Frank kicked five or more goals six times in the first seven matches to lead the VFL goalkicking table. Injury curtailed further progress.

In 1976, Bourke was appointed captain of the club and he responded with a great season, although the team slipped down the ladder. He finished third in the Brownlow medal and third in the club's best and fairest award, demonstrating a phenomenal consistency; in nine seasons between 1968 and 1976, Bourke was placed seven times in the best and fairest count.  However, the first signs of advancing age showed the following year. Bourke was so disappointed with his form at one point he privately contemplated retirement, but he continued on and proved his mettle by captaining Victoria against West Australia and leading Richmond into the finals. Key position defenders were in short supply at Richmond, so Bourke was forced to play at centre half back even though he lacked height for the position. At the end of the season, he decided to resign the captaincy in an effort to prolong his career.  This decision paid off in 1980, when the Tigers returned to power and again won the premiership. Due to his slowing leg speed, Bourke was now at full back but still a formidable opponent. Two weeks before the finals, Bourke entered football folklore in a match at Arden Street against North Melbourne. Bourke received an accidental finger in the eye, which quickly filled with blood that poured down his face and on to his guernsey. At the time there was no "blood rule" (requiring bleeding players to leave the field until the bleeding is stopped) and Bourke shifted to the forward line. Although he could barely see through the mass of blood, he dived full-length to take a chest mark, then kicked an important goal to ensure Richmond had a narrow win.  Bourke, along with Kevin Bartlett, played in all five Richmond premierships of the era, which constitutes the club record. Aged 34, he decided to soldier on in 1981 and became only the third Tiger to play 300 games. However, his form was not always equal to the personal standard that he had set and when it became obvious that Richmond wouldn't make the finals to defend the premiership, Bourke retired. Note that changes to the way records are compiled was made after Bourke's retirement, and his official games tally has been reduced to an even 300, of which 23 were finals and six Grand Finals.
Francis Bourke