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Winchell was born Paul Wilchinsky in New York City on December 21, 1922, to Solomon Wilchinsky and Clara Fuchs. His father was a tailor; his grandparents were Jewish immigrants from Russian Poland and Austria-Hungary. Winchell's initial ambition was to become a doctor, but the Depression wiped out any chance of his family's ability to afford medical school tuition. At age 13, he contracted polio; while recovering, he happened upon a magazine advertisement offering a ventriloquism kit for ten cents.

Winchell's best-known ventriloquist dummies were Jerry Mahoney and Knucklehead Smiff. Mahoney was carved by Chicago-based figure maker Frank Marshall. Sometime later Winchell had basswood copies of Jerry's head made by a commercial duplicating service. One became the upgraded Jerry Mahoney that is seen primarily throughout Winchell's television career. The television versions of Jerry and Knucklehead also featured Winchell's innovation of actors slipping their hands into the sleeves of the dummies, giving the visual effect of gesturing with their hands while "conversing" with each other. He modified two other copies to create Knucklehead Smiff. The original Marshall Jerry Mahoney and one copy of Knucklehead Smiff are in storage at the Smithsonian Institution. The other two figures are in the collection of illusionist David Copperfield.  Winchell's first show as a ventriloquist was on radio with Jerry Mahoney in 1943. The program was short-lived, however, as he was overshadowed by Edgar Bergen. Winchell also created Ozwald, a character that resembled Humpty Dumpty. The effect was accomplished by painting eyes and a nose on his chin, then adding a "body" covering the rest of his face, and finally electronically turning the camera image upside down. In 1961, Berwin Novelties introduced a home version of the character that included an Oswald body, creative pencils to draw the eyes and nose and a "magic mirror" that automatically turned a reflection upside down.  In 1948, Winchell and Joseph Dunninger were featured on Floor Show on NBC. Recorded via kinescope and replayed on WNBQ-TV in Chicago, Illinois, the 8:30-9 p.m. Central Time show on Thursdays was the station's first mid-week program.  During the 1950s, Winchell hosted children's and adult programs with his figures for NBC Television, and later for syndication. The NBC Saturday morning program, sponsored by Tootsie Roll, featured a clubhouse motif and a theme song co-written by Winchell and his longtime bandleader and on-air sidekick, Milton DeLugg. The theme song was entitled "HOORAY, HOORAH" which featured the secret password "SCOLLY WALLY DOO DOO". An ending song entitled "Friends, Friends, Friends" was sung by the children in the audience. On one episode, The Three Stooges appeared on the show to promote their joint feature film venture, Stop, Look, and Laugh, in late 1959. He made an appearance on Nanny and the Professor (Season 2, Episode 13) as a "mean old man" (a puppeteer who had retired into seclusion after losing his wife in an accident). In 1996, Winchell contracted with figure maker Tim Selberg to construct a more contemporary version of Jerry Mahoney, which Winch described as "Disney-esque". Winchell used the new figure version to pitch a new TV series idea to Michael Eisner. In 2009 Winchell was featured in the comedy documentary I'm No Dummy, directed by Bryan W. Simon.
Paul Winchell